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IB DP Biology HL D4.1 Natural selection Flashcards

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[h] IB DP Biology HL D4.1 Natural selection Flashcards

 

[q] D 4.1.1 Natural selection as the mechanism driving evolutionary change

[a] Natural selection – the mechanisn that drives a species to evolve into another species or for a species to remain the same species but still evolve (changes in the species just occur). Aka evolutaionary change.


– organisms that are more adapted to their environment are more likely to survive and pass on the genes that aided their success.


Darwin’s theory of evolution is based on natural selection.


Evolutionary change – the heritable change in populations and species over time, due to mechanisms such as natural selection, random genetic drift, and sexual selection.

 

[q] D 4.1.2 Roles of mutation and sexual reproduction in generating the variations on which natural selection acts

[a] The only way that there are variations to be selected from or against is for there to be mutations and sexual reproduction.


Variations are generated by:


– mutations creating different alleles.


– and sexual reproduction mixing up the different alleles and passing them on.


If ever individual of a species was identical there can not be any natural selection or evolutionary.

 

[q] D 4.1.3 Overproduction of offspring and competition for resources as factors that promote natural selection

[a] Theory of evolution by natural selection depends on:


– more offspring produced than what a environment can support


– and that those organisms have to compete for resources

Carrying capacity – how many of a species an environment can support.

– based on all the biotic and abiotic factors that control how many of a species can live in an environment.

– example of resources: predators, food, nesting, etc.

If there is nothing limitimg a population (it is smaller than what the environment can support) then having one characteristic over another will not make a organism be selected.

 

[q] D 4.1.4 Abiotic factors as selective pressures

[a] Abiotic factors of an ecosystem that can provide selective pressure:


Density independent:


– high/low temperatures


– natural disasters


– the number of the species present does not matter.


Density dependent:


– based on how many of the species is present.

 

[q] D 4.1.5 Differences between individuals in adaptations, survival, and reproduction as the basis for natural selection

[a] Individuals that are best adapted have the highest probability of survival, thus are able to pass on their genes.


Intraspecific competition – competition between individuals of the same species.


Survival value/fitness – a measure of how well a certain phenotype is in a population.


Reproductive potential of a genotype:


– some have a larger potential than others.


– based on the phenotype that the genotype creates.

 

[q] D 4.1.6 Requirement that traits are heritable for evolutionary change to occur

[a] A trait has to be heritable for evolutionary change to occur.


– This is where Lamarck was wrong.


– If a trait is not heritable, it can not be a part of natural selection.

 

[q] D 4.1.7 Sexual selection as a selection pressure in animal species

[a] Sexual selection – a form of natural selection in which the reproductive success of an individual results in more offspring compared to others in the population who do not have as much success in finding a mate. 

Individuals choose who to mate with.


In animal species, sexual selection is a selection pressure that drives the evolution of a population.
Physical/behavioral traits attract animals to one another.


– these traits are signs of fitness.


– example: in a lot of bird species the males are very colorful (the most colorful males are the ones who reproduce the most/are choosen as a mate the most);

females are attracted to the fact that they have been able to survive for so long while having very bright/flashy colors – overall ability to survive is good.

 

[q] D 4.1.8 Modeling of sexual and natural selection based on experimental control of selective pressure

[a] Endler’s guppy experiment (case study of natural selection in the color patterns in guppies)
– wanted to see if the presence of predators modified the bright colors seen in guppies.


– carried out in two ways: in the field (in the streams of Trinidad) and in 10 artificial ponds in a greenhouse at Prinnceton University, USA. (where more variables could be cntrolled than in nature).


– in both the field and in the greenhouse, some populations of guppies where kept with predators while others were not.


– hypothesis: guppies in pools protected from predatory fish would be ornate colors (because there is no threat of their survival);

while guppies in pools where predators where presnt would be less colorful (because there ability to hide would lead to better chances of survival).


– independent variable: presence of predators (one of the selective pressures on male guppies)

 

[q] D 4.1.9 Concepts of the gene pool

[a] Gene pool – all the genetic information present in the reproducing memebrs of a population at a given time.


– all the possible genes (and their alleles) in an interbreeding population.


Inbreeding – when closely related organisms mate wit each other.


– inbreeding narrows the gene pool.


Size of gene pool:


– a population that shows substantial variety in its traits has a larger gene pool.


– a population whose memebers show little variation has a small gene pool.

 

[q] D 4.1.10 Allele frequencies of geographically isolated populations

[a] Allele frequency – a measure of the proportion of a specific version of a gene in a population.


– expressed as a proportion or a percentage.


Geographically isolated populations can have different allele frequencies.


– they are not interbreeding, therefore mutations, differential rates, and etc. can occur.


– example of isolated populations having different gene ratios: ABO blood typing amongst different populations of humans.

 

[q] D 4.1.11 Changes in allele frequency in the gene pool as a consequence of natural selection between individuals according to differences in their heritable traits

[a] Allele frequencies can change as a result of natural selection.


– if one allele has an advantage then it can become a higher frequency in the population.


allele = heritable trait

 

[q] D 4.1.12 Differences between directional, disruptive, and stabilizing selection

[a] Populations of members of the same species (and thus the same gene pool) can be stopped from reprodcuing together because there is an insurmountable barrier between them.


– barriers can be geographical, temporal, behavioural or related to the infertility caused by hybridization.


Directional selection – takes place when one extreme phenotype is favoured over all other phenotypes by natural selection.


– the frequency of one phenotype is seen to increase over time, whereas the other phenotype decreases.


– example: if a drought occurs and the plants that make the little seeds that finches who have dainty bills can eat die off, then the finches whose bills can break the hard seeds will survive and the finches with dainty bills will die off.

 

[q] D 4.1.12 Differences between directional, disruptive, and stabilizing selection (continued)

[a] Disruptive selection – when two extreme phenotypes are favoured by natural selection, rather than one intermediate phenotype.


– can be an advantage (ex. could provide a better chance of survival in places where food sources are vaiable).


– if the differences caused are too extreme and the two populations occupy different niches, it is possible for speciation to occur.


– example: in a species of invertebrate that has body shades that go from very light to very dark, both the very light and very dark will survive by living on different parts of the sandy beach that best fit their coloring (along the ocean floor or within the rocks).


Stabilizing selection – when one phenotype is favoured over two extreme phenotypes.


– when the average is selected.


– example: birth weight in humans. Babies that are born too small can lose heat too easily and may die, whereas babies being born too large can lead to complications during childbirth and the death of the mother or the baby.


All of these types of selection change allele frequencies.


– any alleles that do not led to the favoured phenotype(s) will not be selected for and will make up less of the next generation.


– any of the alleles that are favoured will be more popular.

 

[q] D 4.1.13 Hardy-Weinberg equation and calculations of allele or genotype frequencies

[a] Hardy-Weinberg equation and calculations of allele or genotype frequencies:


p + q = 1 (frequency of the alleles)


p^2 + 2pq + q^2 = 1 (frequency of the genotypes)


Dominant allele = p


Recessive allele = q


homozygous dominant = p^2


heterozygous = 2pq


homozygous recessive = q^2


If you know the frequency of the HOMOZYGOUS RECESSIVE, you can figure out the frequency of the dominant allele.


Examples:


– freckles


– PTC tasting

 

[q] D 4.1.14 Hardy-Weinberg conditions that must be maintained for a population to be in genetic equilibrium

[a] Hardy-Weinberg conditions that must be maintained for a population to be in genetic equilibrium:


– No gene mutations


– Mating is random


– No immigration or emigration


– Population large enough for chance events to alter frequencies


– Natural selection does not favor one phenotype


*If an allele frequency is calculated based on Hardy-Weinberg then compared to previous calculations and the frequency is not the same/has to had changed, one of these conditions had to have not been maitained.


Genetic equilibrium – the condition where a gene pool is not changing in frequency across generations.

 

[q] D 4.1.15 Artificial selection by deliberate choice of traits

[a] Artificial selection – the process by which humans breed organisms to increase what we see as desirable characteristics.


– is performed by selective breeding.


– when specific traits are slected for.


Artificial selection leds to an artificial change in allele frequencies.

 

[q] Linking Questions

[a] 1. How do intraspecific interactions differ from interspecific interactions?


2. What mechanisms minimize competition?

 

[q] Variation

[a] Natural Phenomenon. 

Not always seen, as variation can be genotypic, or phenotypic

Causes: 

– Mutation, Any change to the DNA sequence 

-Meiosis, Produce gametes containing unique combinations of alleles, increasing genetic variation of a species

-Sexual Reproduction, Combination of gametes results in a zygote with even more increased variation

 

[q] Adaptation 

[a] Unconscious adaptations made populations rather than individuals, as if their characteristic fits, they will have a better life.

 

[q] Overproduction of Offspring 

[a] Rare phenomenon in which parents produce too many offspring, meaning that those with the best features will survive, and be more likely to reproduce, creating natural selection

 

[q] Inheritance

[a] Individuals that do reproduce pass on characteristics to their offspring.


Natural selection therefore increases the frequency of favourable characteristics in a population, meaning individuals will be better adapted, results in a change of species

 

[q] Process – Natural Selection

[a] 1. Variation exists within any population of organisms 

2.Organisms with suitable features for environment will survive and reproduce, whereas others will not. 

3. Organisms that are able to survive can reproduce and pass on their heritable characteristics to their population 

4. Population will become more suited to the environment 

5. Natural selection acts to increase the frequency of characteristics that make individuals better adapted 

 

[q] Differential Survival and Reproduction

[a] Individuals are better adapted tend to survive and reproduce, while the less adapted tend to die or produce fewer offspring.

[q] Progressive Change

[a] Natural selection increases the frequency of characteristics that make individuals better adapted and decreases the frequency of other characteristics leading to changes within the species

[q] Galapagos Finches

[a] 14 different species unevenly distributed in Galapagos Islands


They evolved to adapt to particular food source, with large beaks appearing for large seeds, and small beaks appearing for small seeds, which allowed the various finch populations to survive side by side.

[q] Geospiza Fortis

[a] Could live off both small and large seeds because of variation in beak size.

1977, a drought occurs, causing a shortage smaller seeds, and the birds with longer and narrower beaks survived, as they could survive on large seeds.

El Nino of 1982/83 caused massive rainfall, meaning the G.fortis population increased due to ample small seed availability.

With the return of the drought in 1987, only a small cohort of birds were still breeding, but they were not a random sample of the earlier Geospiza population.

[q] Antibiotic resistance 

[a] Antibiotics provide a selection pressure and because of the variation in a population.

Some bacteria will be resistant to the antibiotic, meaning that if the resistance is heritable, it will pass into future generations, as well as other cells via plasmid transfer.


Bacteria may also gain resistance through a misuse of antibiotics, as they have been exposed, and can develop resistance.

[q] D4.1.1—Natural selection as the mechanism driving evolutionary change 
Students should appreciate that natural selection operates continuously and over billions of years, resulting in the biodiversity of life on Earth.
[a]  Natural selection is the process by which better adapted organisms with favorable alleles survive and pass their genes to offspring, taking over the population’s gene pool and weeding out deleterious ones to produce the fitter, not fittest (since it operates continuously).
It is the main mechanism driving evolutionary change.
Natural selection benefits the individual and not necessarily the species.
A strong group of lions may kill a weaker group and breed with the females.
Although the total number of individuals within the species has momentarily decreased, genes from stronger lions increased in frequency to allow for more adapted offspring.
[q] D4.1.6—Requirement that traits are heritable for evolutionary change to occur 
Students should understand that characteristics acquired during an individual’s life due to environmental factors are not encoded in the base sequence of genes and so are not heritable.
[a] Only heritable traits (those that are encoded within an individual’s genome) are able to be passed down and be a part of an evolutionary change due to natural selection.
Characteristics acquired during an individual’s life due to environmental factors, such as certain behaviors or knowledge, are not part of the genome and thus are not heritable.
Additional higher level: although epigenetic tags could be inherited in gametes and thus passed to offspring (gene imprinting), they do not change DNA sequences and thus far are not regarded as having a role in evolutionary change
[q] D4.1.5—Differences between individuals in adaptation, survival and reproduction as the basis for natural selection
Students are required to study natural selection due to intraspecific competition, including the concept of fitness when discussing the survival value and reproductive potential of a genotype. 
[a] Intraspecific competition occurs between individuals of the same species.
‘Fitness’ is how well-adapted an individual is to their environment; the fitter they are the greater their survival value and reproductive potential.
Thus, they will contribute the most to the gene pool of the species (their alleles will spread the most within the species) and evolutionary change will occur.
[q] NOS: In Darwin’s time it was widely understood that species evolved, but the mechanism was not clear.
Darwin’s theory provided a convincing mechanism and replaced Lamarckism.
This is an example of a paradigm shift.
Students should understand the meaning of the term “paradigm shift”.
[a] A paradigm shift is a fundamental change in the basic concepts and approaches in a scientific discipline.
Lamarckism was a previously accepted evolutionary theory until Darwinism came and providing more a more convincing mechanism of evolution, leading to a paradigm shift in biology
[q] D4.1.2—Roles of mutation and sexual reproduction in generating the variation on which natural selection acts 
Mutation generates new alleles and sexual reproduction generates new combinations of alleles. 
[a] Variation within species provides different alleles of the same gene which allows natural selection to decrease or increase allele frequency already within the population.
If there is no variation, all individuals have the same reproductive and survival success rate because they all have the same traits, thus natural selection will not operate because no alleles of which to choose from even exist.
Mutations drive the genetic variation which natural selection acts on.
Although mutations are random in the sense that they occur regardless of whether they are harmful or beneficial, natural selection is not random as it only chooses the beneficial mutations that result in better adaptations.
These mutations produce new alleles by single point mutations, insertions, and deletions, among others.
Sexual reproduction also contributes to variation within a species through crossing over in prophase I of meiosis and random/independent assortment of homologous pairs in metaphase I.
[q] D4.1.3—Overproduction of offspring and competition for resources as factors that promote natural selection
Include examples of food and other resources that may limit carrying capacity.
[a] In nature, population sizes are limited by many factors, some of which are density-dependent or density-independent.
These factors act as selection pressures, which are biotic or abiotic agents that affect the ability of an organism to reproduce and survive.
Density-dependent factors arise from biological phenomena like competition between populations, competition, disease, and predation.
They arise due to interactions between organisms, and can either increase or decrease population size and mortality through changes in reproduction and survival, driving natural selection.
These factors usually serve to regulate population size and keep it within a narrow range.
For instance, species have the tendency to overproduce offspring beyond what the environment can provide in terms of resources.
This leads to competition between individuals of the same species, which promotes natural selection as only those with favorable alleles can successfully compete and utilize the limited resources.
Thus, natural selection acted due to the presence of a density-dependent factor
[q] D4.1.4—Abiotic factors as selection pressures 
Include examples of density-independent factors such as high or low temperatures that may affect survival of individuals in a population.
[a] Density-independent factors arise from physical or chemical phenomena (abiotic factors) like natural disasters, weather conditions, quality and quantity of food, and pollution.
For example, a rise in temperature may kill some plants within a species, but those who survive are better adapted and thus able to reproduce.
Thus, natural selection selects those adapted to hotter temperatures.
[q] D4.1.7—Sexual selection as a selection pressure in animal species 
Differences in physical and behavioural traits, which can be used as signs of overall fitness, can affect success in attracting a mate and so drive the evolution of an animal population.
Illustrate this using suitable examples such as the evolution of the plumage of birds of paradise.
[a] Sexual selection is a subset of natural selection that allows sexually selected traits to evolve if they offset the male’s diminished survival with an increase in his reproduction.
Females prefer males with stronger traits (disease resistance, size, etc.), and certain aesthetic features that serve as indicators for these strong traits (like brighter colors).
For many species, these traits do not help the organism adapt to its environment, and expediate quite a lot of energy despite natural selection being responsible for their development.
The whole process of sexual selection stops only when the male trait becomes so exaggerated that any further increase reduces his survival more than it attracts females, so that his net production of offspring suffers.
For example, colorful plumage in male birds of paradise have evolved to attract female mates by indicating desirable traits in the male.
An experiment was done to add a white aesthetic feature to the plumage of male birds, and it was observed that females were heavily attracted to it and mated more with males who possess the added white feature.
A possible explanation by scientists posits that female birds line their nests with white feathers, so the white addition on the male plumage indicated to the females that they are mating with a male who can take care of their offspring after birth.
In any case, evolutionary theory shows that in events of sexual selection, three types of genes will all increase in frequency together: genes for a male “indicator” trait reflecting that he has good genes, genes that make a female prefer that indicator trait, and the “good” genes in males whose presence is reflected by the indicator.
[q] D4.1.8—Modelling of sexual and natural selection based on experimental control of selection pressures
Application of skills: Students should interpret data from John Endler’s experiments with guppies.
[a] John Endler investigated sexual selection through guppies, a type of fish.
Male guppies living in environments with dangerous predators were less colorful than those living in safer waters as they camouflaged better, even though females prefer those with more colors.
He conducted modelling experiments by transferring male guppies living in dangerous waters to safer areas, and after 2 years (around 15 generations) sexual selection favored those with brighter colors.
[q] D4.1.9—Concept of the gene pool 
A gene pool consists of all the genes and their different alleles, present in a population. 
[a] A gene pool consists of all the genes and their different alleles present in a population or species.
[q] D4.1.10—Allele frequencies of geographically isolated populations 
Application of skills: Students should use databases to search allele frequencies.
Use at least one human example.
[a] Species may exist in different geographically isolated environments and thus individuals may belong to different populations.
Despite belonging to the same species, these populations have distinct gene pools with different allele frequencies.
For example, the lactose tolerance allele in humans exists in different frequencies across the many human populations on earth (people living in certain continents or regions are more likely to be lactose intolerant than others).
[q] D4.1.11—Changes in allele frequency in the gene pool as a consequence of natural selection between individuals according to differences in their heritable traits
Darwin developed the theory of evolution by natural selection.
Biologists subsequently integrated genetics with natural selection in what is now known as neo-Darwinism
[a] Darwin developed the theory of evolution by natural selection but was unable to fully explain all its details (DNA was not yet even known back then), but biologists subsequently integrated new discoveries in genetics with natural selection to emerge with what is now known as neo-Darwinism.
Essentially, natural selection increases the frequency of beneficial alleles and reduces that of harmful ones within the gene pool.
[q] D4.1.12—Differences between directional, disruptive and stabilizing selection 
Students should be aware that all three types result in a change in allele frequency. 
[a] Natural selection on traits controlled by a single gene changes frequency of its alleles, but with polygenic traits (characteristics controlled by multiple genes), natural selection affects the distribution of phenotypes in three main ways:
1. Stabilizing selection: the intermediate phenotype is chosen as the population stabilizes on a single trait, which decreases genetic diversity due to homogeneity.
o (i.e. human birth weight is a type of stabilizing selection since large babies caused pregnancy complications and light babies were less likely to survive, so the phenotype stabilized in the middle at around 7lb)
2. Directional selection: one of two phenotypic extremes within the spectrum is selected for, causing the allele frequency to continuously shift towards one ‘direction’.
This usually occurs in highly changeable environments or when a new allele develops (due to mutation) and overpowers all others due to its benefits.
o (i.e. during the industrial revolution, air pollution caused tree bark to turn into a black color, so black peppered moths were more likely to survive than white ones due to camouflage)
3. Disruptive (diversifying) selection: when two extreme/divergent phenotypes are favored over intermediate ones, increasing genetic diversity and heterogeneity.
o (i.e. big male elephants can mate by brute force and small ones can sneak into the big males’ territory and mate whereas medium-sized elephants are too big to sneak in and too small to overtake the large elephants in a battle)
[q] D4.1.14—Hardy–Weinberg conditions that must be maintained for a population to be in genetic equilibrium
Students should understand that if genotype frequencies in a population do not fit the Hardy–Weinberg equation, this indicates that one or more of the conditions is not being met, for example mating is non-random or survival rates vary between genotypes.
[a] Genetic equilibrium is a condition in which the allele frequency within a gene pool does not change across generations, satisfying the following conditions:
• no mutations
• no gene flow (individuals do not enter or exit the population)
• random mating (individuals do not favor some characteristics over others)
• no natural selection
This state of equilibrium can be attributed either to the entire genome or to one specific allele.
If we look at the entire genome, then these conditions have to be met for every allele, but we look at only one single allele then these criteria have to be met for only that one single allele.
[q] D4.1.14—Hardy–Weinberg conditions that must be maintained for a population to be in genetic equilibrium
Students should understand that if genotype frequencies in a population do not fit the Hardy– Weinberg equation, this indicates that one or more of the conditions is not being met, for example mating is non-random or survival rates vary between genotypes.
[a] The Hardy-Weinberg Principle of Equilibrium states that allele frequencies are inherently stable as long as no evolutionary force is acting on them.
While some genes may be at equilibrium, is it highly unlikely (almost impossible) for a population to have all of its genes at equilibrium.
Even though no population on earth satisfies the Hardy-Weinberg equilibrium, it provides a baseline to which we can compare actual populations to in order to detect how evolutionary forces are acting on the population’s gene pool.
For example, when conducting a chi-square test, expected frequencies at equilibrium can be compared to actual observed frequences to quantify the degree to which the population’s alleles are deviated away from equilibrium.
[q] D4.1.13—Hardy–Weinberg equation and calculations of allele or genotype frequencies 
Use p and q to denote the two allele frequencies. Students should understand that p + q = 1 so genotype frequencies are predicted by the Hardy–Weinberg equation: p2 + 2pq+ q2 = 1.
If one of the genotype frequencies is known, the allele frequencies can be calculated using the same equations.
[a] Suppose a gene has two alleles only, p and q. The allele frequencies of both p and q add up to 1 (100%).
If an individual is homozygous for the allele, the allele is counted twice (number of individuals with allele ¹ allele frequency).
Not only does allele frequency interest biologists, but genetic frequency as well, which describes the frequency of a specific genotype (combination of alleles i.e. homozygous pp/qq or heterozygous pq) within a population.
This can be calculated through the following equation:
• p2 = frequency of individuals who are pp homozygous
• 2pq = frequency of individuals who are pq heterozygous
• q2 = frequency of individuals who are qq homozygous
[q] D4.1.15—Artificial selection by deliberate choice of traits 
Artificial selection is carried out in crop plants and domesticated animals by choosing individuals for breeding that have desirable traits.
Unintended consequences of human actions, such as the evolution of resistance in bacteria when an antibiotic is used, are due to natural rather than artificial selection.
[a] Artificial selection is essentially natural selection but instead of selecting the traits that make the species more adapted to its environment, humans deliberately select for traits most beneficial to them (i.e. domesticating faster horses or more weather-resistant crops).
This has been used as evidence for evolution as it can show the gradual evolutionary change much faster than what is observed in nature.
 
 

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